Gramma, a 141-year-old Galapagos tortoise and the San Diego Zoo’s oldest resident, has died. She came from the Bronx Zoo nearly a century ago and was known as the "Queen of the Zoo." Visitors and ...
LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.
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